NYPD detective Danny Reagan teams up with Boston detective Lena Silver to investigate a deadly fire at a tech company. As they dig deeper, they uncover a complex web of secrets involving facial recognition technology, family ties and a search for justice that brings the Silver and Reagan families closer together.

Lena and Danny are pulled into a high-stakes cuddle investigation when a whistleblower tied to one of Mae's court cases is found dead. Meanwhile, Jonah and Sean prepare for their first official day on the job.

A body found on the Boston wharf pulls Lena and Danny into a tense drug investigation alongside Lena's former partner, Detective Brian Rodgers, sparking personal and professional friction. Jonah and Sean chase down a missing man tied to a scavenger hunt, while Sarah and Mae navigate emotional challenges at home and in court.

As the Silver family observes Yahrzeit to honor patriarch Ben Silver on the one-year anniversary of his passing, Lena and Danny investigate the cuddle of a beloved shop owner. Meanwhile, Sarah responds to a hostage crisis and Mae faces a difficult legal decision.