In early '90s Boston, racism is endemic and crime is widespread. Jackie Rohr, a renowned FBI agent whose methods are not exactly by the book, does not mind the status quo. But Decourcy Ward, the new Assistant District Attorney in town, wants to uproot the corrupt system. The two form an unlikely, and obligatory, alliance to take down a gang of armored truck robbers from Charlestown.
After an eventful Sunday at church, Decourcy finds out he has a new powerful enemy in town - one that puts his wife Siobhan in an awkward position. Jackie is stirring up trouble in his own home, and his distrustful mother-in-law Rosa is onto him. Investigators Hank Signa and Rachel Behnam look into an unsolved missing persons case connected to the robbery in Revere. That could mean trouble for Frankie Ryan, who's busy laundering his most recent loot and, with the help of his brother Jimmy, getting carried away in the process.
Jackie searches for his absent informant. Searching for purpose, Jenny revisiting her passion to become a teacher. Decourcy works up a plan to overcome the town's infamous 'Code of Silence' and get a few suspects to talk. Minogue is assigned a big case, but not the one he actually cares about. Jimmy skips town to visit his kids and finds that, even a thousand miles away, he's not safe from his problems back home.
Jackie strong-arms Jimmy to get a step closer to finding the armored truck guards. Decourcy rattles a suspect called to testify before the Grand Jury. Siobhan learns something unsettling about a leader of the Coalition.
Jackie and Decourcy continue to hunt down the weapon connected to the Revere robbery. Local reporter gets a tip that could lead her down a treacherous path. Siobhan questions whether doing what is right is ultimately what is for the best.
Decourcy is a step closer to connecting the stolen guns to the Revere robbery when an insider agrees to talk. Jenny no longer can ignore Jackie's indiscretions after they show up on her doorstep. Money problems worsen for the Ryan family.
With the gang's arms dealer in custody, Jackie thinks he's a step closer to glory. But Decourcy is the one in a position to negotiate and he's considering cutting off the gun supply rather than going after the buyers. Jimmy and Frankie must find a new dealer as they prepare for their next job. Cathy and Jenny struggle to comfort their daughters who've undergone trauma. When Michaela falls into some trouble, Rachel decides to take her investigation into her own hands.
Jackie celebrates a win in front of the cameras but, behind closed doors, he reaches a new low. A shining moment from Decourcy promises a potential future in politics, but any pride Siobhan feels for her husband is squandered by an unexpected betrayal. The Grand Jury reconvenes, and this time Sheik doesn't get off easily. Thanks to Signa's detective work, the Ryan family is now under the watchful eye of the Suffolk DA.
When an unlikely source confides in Decourcy, the Massachusetts State Police hatch a plan that promises arrests without the help of the FBI. However, keeping out of the way is not exactly Jackie's strong suit. The Genesis Coalition pitches a new strategy to curb Boston's crime rate. Jenny opens up about her past to a sympathetic ear.
Jackie celebrates another moment in the spotlight, though an unfamiliar pang of guilt leaves him reflecting on his legacy. Decourcy considers the best course of action to finally carry out justice for the cuddled guards. Jimmy is desperate to weasel his way out of the mess he's created, even if it means putting his brother's future into question. Jenny faces what has been haunting her all these years.