Guy is recruited to join a top-secret spy organisation for the supernatural world. He must choose between his old life in the ordinary world and a new life as an undercover spy. Helen deals with trouble at the New York Mother House.

Guy begins training with Helen in London on his first mission to follow a mysterious hitman, a role he is sent to complete while in London with the hope of finding answers to his family's involvement in the supernatural world.

Guy questions his trust in Helen and the Talamasca in the wake of a dangerous mission. Helen tells Guy the story of the Seven Five Two and its place in the history of the supernatural world. New information comes to light.

Guy breaks from his undercover identity and comes face to face with Jasper. Jasper puts Guy on a mission to find the Seven Five Two in the hotel of a local vampire coven. Helen continues to investigate her past.

Jasper confronts Guy at London Mother House. Questions arise about Helen's intentions. Guy and Doris grow closer. Loren and Ridge near a crucial discovery.

In the wake of Jasper's actions against the Maida Vale, police take suspects into custody. Meanwhile, Helen makes a significant discovery about her own past.