Desperate circumstances call for desperate measures, as Jake, Holt, Amy, Terry, Rosa and Charles must ally themselves with old enemies.
Detective Jake Peralta is a good enough cop that he's never had to work that hard or follow the rules too closely. Perhaps because he has the best arrest record among his colleagues, he's been enabled - if not indulged - throughout his entire career. That is, until the precinct gets a new commanding officer, Captain Ray Holt, who reminds this hotshot cop to respect the badge. Jake may have collared more criminals, but Detective Amy Santiago is close behind, and she's keenly aware of how many arrests she needs to close the gap. Amy grew up with seven brothers who were all cops. She's the first girl in the family to put on a police uniform, and suffice it to say: she's extremely competitive... about everything.
Also working cases in Brooklyn's 99th precinct is Sergeant Terry Jeffords, a linebacker of a man who's lost his nerve, not because he's a wimp, but because a year ago, his wife had twin baby girls - Cagney and Lacey - and he can't imagine not seeing them grow up. Detective Charles Boyle, who idolizes Jake, is the precinct's workhorse; he's not that brilliant, he's not physically gifted, but he tries harder than anyone else. Charles pines for Detective Rosa Diaz, with whom he stands no chance at all. Rosa is simultaneously tough, sexy and scary as hell. She's vocally opinionated about everything. Cleaning up everyone's mess is Gina Linetti, the eccentric, civilian office manager who somehow gets involved in everyone's business. Together, these tightly knit - or is it tightly wound? - cops interrogate suspects, arrest perps and solve murders.